The earliest known crown-Testudo tortoise from the late Miocene (Vallesian, 9 Ma) of Greece Garcia, Géraldine ; Pinton, Aurélie ; Valentin, Xavier ; Kostopoulos, Dimitris S. ; Merceron, Gildas ; de Bonis, Louis ; Koufos, George D. Abstract Publication: PLoS ONE Pub Date: April 2020 DOI: 10.1371/journal.pone.0224783 Bibcode: 2020PLoSO..1524783G