Two new polyphagan beetles (Tenebrionidae, Leiodidae) from lower Cenomanian amber of Myanmar

Mesozoic tenebrionid and leiodid beetles are poorly known. Here we describe Praezolodinus pilosus gen. et sp. nov. (Tenebrionidae: Zolodininae) and Cretoptomaphagus microsoma gen. et sp. nov. (Leiodidae: Cholevinae), both from mid-Cretaceous Burmese amber (ca. 99 Ma). Praezolodinus pilosus is placed in the basal group Zolodininae due to (1) galea preserved with fine and long pubescence, (2) aedeagus inverted, pimeloid type. Cretoptomaphagus microsoma is assigned to Leiodidae and included in Cholevinae based on its (1) characteristic interrupted five-segmented antennal club, and (2) cephalic configuration, with the posterodorsal contour of the exposed part of the head abutting with the anterior pronotal edge. Cretoptomaphagus microsoma is the second Cholevinae described from the Mesozoic. Its strong morphological similarity with extant members of Cholevinae makes this fossil an astonishing example of long-term morphological stasis in Coleoptera.