The immense diversity of the galaxy population in the universe is believed to stem from their disparate merging histories, stochastic star formations, and multi-scale influences of filamentary environments. Any single initial condition of the early universe was never expected to explain alone how the galaxies formed and evolved to end up possessing such various traits as they have at the present epoch. However, several observational studies have revealed that the key physical properties of the observed galaxies in the local universe appeared to be regulated by one single factor, the identity of which has been shrouded in mystery up to date. Here, we report on our success of identifying the single regulating factor as the degree of misalignments between the initial tidal field and protogalaxy inertia momentum tensors. The spin parameters, formation epochs, stellar-to-total mass ratios, stellar ages, sizes, colors, metallicities and specific heat energies of the galaxies from the IllustrisTNG suite of hydrodynamic simulations are all found to be almost linearly and strongly dependent on this initial condition, when the differences in galaxy total mass, environmental density and shear are controlled to vanish. The cosmological predispositions, if properly identified, turns out to be much more impactful on galaxy evolution than conventionally thought.