Cycads in the Permian of the Subangara Region

Remains of cladosperms attributed to Cycadales and described as Dioonitocarpidium rossicum sp. nov. were found in the Kazanian of the Russian Platform. They confirm the presence of Cycadales in the Permian of the Subangara Region, previously suggested by finds of vegetative leaves. The significance of Dioonitocarpidium as a possible initial form in the evolution of the Cycadales in the Mesozoic and Cenozoic is discussed.