Crustacea (Anomura, Brachyura) from the Miocene of Veracruz and Chiapas, Mexico: New records and new species

We report several new crab specimens, representing new taxa and new occurrences from the Miocene of Veracruz and Chiapas, SE Mexico. Most of the specimens were collected by the late Emily H. Vokes, in 1963 from outcrops of the Agueguexquite Formation, Veracruz, Mexico. The age range of the studied material support an upper Miocene age for the Agueguexquite Formation, as previously suggested. Ranina sp. and Portunus sp. cf. P. atecuicitlus Vega et al.1999 from the middle Miocene, Tuxpan Formation, are also reported. We also illustrate an incomplete portunoid specimen from the lower Miocene Tulijá Formation of Chiapas, similar to Necronectes A. Milne Edwards, 1881. The presence of Hepatus Latreille, 1802, Persephona Leach, 1817, and Euphylax Stimpson, 1862a, in the Agueguexquite Formation represent the first record of these genera in the fossil record of Mexico. The occurrence of Eupilumnus? vokesae Luque, sp. nov. would represent the first fossil record of the genus known to date, extending its stratigraphic record into the Miocene. Eurytium granulosus Schweitzer, Velez-Juarbe, Martinez, Collmar-Hull, Feldmann and Santos, 2008, first reported from the Miocene of Puerto Rico is also identified among the specimens here studied.