Molecular phylogeny of extant Holothuroidea (Echinodermata)

First multigene phylogeny that spans extant Holothuroidea. A revised taxonomy of Holothuroidea is proposed. Three families are redefined and two new families are erected. Discrepancies between time-tree analyses suggest the need for fossil revision. Mapping morphology reveals homoplasy but also recovers apomorphies for clades.