The major development in this paper is a method for determining rates of water movement within the soil profile on watersheds under natural conditions. The method uses rainfall rates or infiltration curves as the supply of water to the soil and field measurements of soil-moisture changes. The major objective was the determination of relationships between infiltration, soil moisture, and surface runoff on watersheds representing two soil types of different hydrologic characteristics and in three vegetal covers. Information was developed on the effect of soil properties and land use upon such relationships.The amounts of water taken up by successive inch increments of soil depth during storms were established by soil-moisture measurements. The time required for each of these amounts to accumulate was determined from the accumulated infiltration curve, or from rainfall minus interception storage when no runoff occurred. Analyses indicate the velocity of water moving through the soil under various conditions, particularly differences in available storage space. Transmission rates or velocities as high as ten in/hr were obtained for the topsoil or plow layer. For a number of storms, water reached the subsoil shortly after runoff began. After water reached the subsoil, the rates at which the remaining topsoil storage was exhausted were deducted from the infiltration rates and the percolation rates at the bottom of the topsoil established. Percolation rates did not exceed 0.60 in/hr. Using percolation rates as the supply, transmission rates through the subsoil were established. Transmission rates did not exceed 1.50 in/hr for the subsoil. As saturation was approached, rates of infiltration decreased until they approached a constant rate at saturation and the low transmission rates in the subsoil became the controlling factor. Infiltration curves, percolation curves, and transmission rates are illustrated and findings interpreted in the light of soil properties and land use.